ARADHYA M K, POTTER D, GAO F Y, et al. Molecular phylogeny of Juglans (Juglandaceae): A biogeographic perspective[J]. Tree Genetics & Genomes, 2007, 3(4): 363378.

ZHAO P, ZHOU H J, POTTER D, et al. Population genetics, phylogenomics and hybrid speciation of Juglans in China determined from whole chloroplast genomes, transcriptomes, and genotyping-by-sequencing (GBS) [J]. Molecular Phylogenetics and Evolution, 2018, 126: 250-265.

FENG X J, ZHOU H J, ZULFIQAR S, et al. The phytogeographic history of common walnut in China[J]. Frontiers in Plant Science, 2018, 9: 1399.

MANNING W E. The classification within the Juglandaceae[J]. Annals of the Missouri Botanical Garden, 1978, 65(4): 1058-1087.

MANOS P S, STONE D E. Evolution, phylogeny, and systematics of the Juglandaceae[J]. Annals of the Missouri Botanical Garden, 2001, 88(2): 231-269.

RORABAUGH J M. English and black walnut phenolic antioxidant activity in vitro and following human nut consumption [J]. Food and Nutrition Sciences, 2011, 2(3): 193-200.

ELOUAFY Y, EL IDRISSI Z L, EL YADINI A, et al. Variations in antioxidant capacity, oxidative stability, and physicochemical quality parameters of walnut (Juglans regia) oil with roasting and accelerated storage conditions[J]. Molecules, 2022, 27(22): 7693.

GAO P, LIU R J, JIN Q Z, et al. Comparative study of chemical compositions and antioxidant capacities of oils obtained from two species of walnut: Juglans regia and Juglans sigillata[J]. Food Chemistry, 2019, 279: 279-287.

ROS E, MATAIX J. Fatty acid composition of nuts: Implications for cardiovascular health[J]. The British Journal of Nutrition, 2006, 96(S2): S29-S35.

TORABIAN S, HADDAD E, CORDERO-MACINTYRE Z, et al. Long-term walnut supplementation without dietary advice induces favorable serum lipid changes in free-living individuals[J]. European Journal of Clinical Nutrition, 2010, 64(3): 274-279.

VINSON J A, CAI Y X. Nuts, especially walnuts, have both antioxidant quantity and efficacy and exhibit significant potential health benefits[J]. Food & Function, 2012, 3(2): 134-140.

奚声珂. 我国胡桃属(Juglans L.)种质资源与核桃(Juglans. regia L.)育种 [J]. 林业科学, 1987, 23(3): 342-350.

XI S K. Gene resources of Juglans and genetic improvement of Juglans regia in China [J]. Scientia Silvae Sinicae, 1987, 23(3): 342-350.

郗荣庭. 中国核桃(Juglans regia L.)起源考证[J]. 河北农业大学学报, 1990, 13(1): 89-94.

XI R T. Textural criticism of walnut (Juglans regia L.) origin in China[J]. Journal of Hebei Agricultural University, 1990, 13(1): 89-94.

路安民, 张志耘. 胡桃目的分化，进化和系统关系[J]. 植物分类学报, 1990, 28(2): 96-102.

LU A M, ZHANG Z Y. The differentiation, evolution and systematic relationship of Juglandales[J]. Acta Phytotaxonomica Sinica, 1990, 28(2): 96-102.

路安民. 论胡桃科植物的地理分布[J]. 植物分类学报, 1982, 20(3): 257-274.

LU A M. On the geographic distribution of the Juglandaceae[J]. Acta Phytotaxonomica Sinica, 1982, 20(3): 257-274.

赵鹏, 周惠娟, 刘占林, 等. 胡桃属植物分子系统发育和生物地理研究进展[J]. 林业科学, 2014, 50(11): 147-157.

ZHAO P, ZHOU H J, LIU Z L, et al. A review of research progress on molecular phylogeny and biogeography in Juglans[J]. Scientia Silvae Sinicae, 2014, 50(11): 147-157.

ZHAO P, ZHAO G F, ZHANG S X, et al. RAPD derived markers for separating Manchurian walnut (Juglans mandshurica) and Japanese walnut (J. ailantifolia) from close congeners [J]. Journal of Systematics and Evolution, 2014, 52(1): 101-111.

ZHAO P, WOESTE K E. DNA markers identify hybrids between butternut (Juglans cinerea L.) and Japanese walnut (Juglans ailantifolia Carr.)[J]. Tree Genetics & Genomes, 2011, 7(3): 511533.

ZHAO P, WOESTE K E, ZHANG S X. Molecular identification and genetic analysis of Juglans resources[M]. Saarbrücken: Lambert Aacdemic Publishing, 2012: 3-20.

ZHAO P, ZHANG S X, WOESTE K. Genotypic data changes family rank for growth and quality traits in a black walnut (Juglans nigra L.) progeny test[J]. New Forests, 2013, 44(3): 357-368.

胡昳恒, 党萌, 张甜, 等. 秦岭地区核桃自然群体和栽培群体的遗传多样性及其演化关系：基于nrDNA ITS序列分析[J]. 林业科学, 2014, 50(12): 47-55.

HU Y H, DANG M, ZHANG T, et al. Genetic diversity and evolutionary relationship of Juglans regia wild and domesticated populations in Qinling mountains based on nrDNA ITS sequences [J]. Scientia Silvae Sinicae, 2014, 50(12): 47-55.

张甜, 王玛丽, 赵鹏. 基于核基因序列JRD5680的核桃群体遗传多样性和遗传结构研究 [J]. 植物研究, 2016, 36(2): 232-241.

ZHANG T, WANG M L, ZHAO P. Sequence analysis of nuclear DNA JRD5680 for determining genetic diversity and genetic structure analysis of common walnut (Juglans regia L.) [J]. Bulletin of Botanical Research, 2016, 36(2): 232-241.

HU Z, ZHANG T, GAO X X, et al. De novo assembly and characterization of the leaf, bud, and fruit transcriptome from the vulnerable tree Juglans mandshurica for the development of 20 new microsatellite markers using Illumina sequencing[J]. Molecular Genetics and Genomics: MGG, 2016, 291(2): 849-862.

BAI W N, LIAO W J, ZHANG D Y. Nuclear and chloroplast DNA phylogeography reveal two refuge areas with asymmetrical gene flow in a temperate walnut tree from East Asia[J]. The New Phytologist, 2010, 188(3): 892-901.

POTTER D, GAO F Y, BAGGETT S, et al. Defining the sources of Paradox: DNA sequence markers for North American walnut (Juglans L.) species and hybrids[J]. Scientia Horticulturae, 2002, 94(1/2): 157-170.

POLLEGIONI P, WOESTE K, OLIMPIERI I, et al. Long-term human impacts on genetic structure of Italian walnut inferred by SSR markers[J]. Tree Genetics & Genomes, 2011, 7(4): 707-723.

MALVOLTI M E, FINESCHI S, PIGLIUCCI M. Morphological integration and genetic variability in Juglans regia L.[J]. Journal of Heredity, 1994, 85(5): 389-394.

STANFORD A M, HARDEN R, PARKS C R. Phylogeny and biogeography of Juglans (Juglandaceae) based on matK and ITS sequence data[J]. American Journal of Botany, 2000, 87(6): 872-882.

FJELLSTROM R G, PARFITT D E. Phylogenetic analysis and evolution of the genus Juglans (Juglandaceae) as determined from nuclear genome RFLPs [J]. Plant Systematics and Evolution, 1995, 197(1): 19-32.

HU Y H, DANG M, FENG X J, et al. Genetic diversity and population structure in the narrow endemic Chinese walnut Juglans hopeiensis Hu: Implications for conservation[J]. Tree Genetics & Genomes, 2017, 13(4): 91.

HU Y H, WOESTE K E, ZHAO P. Completion of the chloroplast genomes of five Chinese Juglans and their contribution to chloroplast phylogeny[J]. Frontiers in Plant Science, 2017, 7: 1955.

FENG X J, YUAN X Y, SUN Y W, et al. Resources for studies of iron walnut (Juglans sigillata) gene expression, genetic diversity, and evolution[J]. Tree Genetics & Genomes, 2018, 14(4): 51.

MU X Y, SUN M, YANG P F, et al. Unveiling the identity of wenwan walnuts and phylogenetic relationships of Asian Juglans species using restriction site-associated DNA-sequencing[J]. Frontiers in Plant Science, 2017, 8: 1708.

MANCHESTER S R. The fossil history of the Juglandaceae[M]. Saint Louis: Missouri Botanical Garden, 1987: 1-37.

吴燕民, 裴东, 奚声珂, 等. 用RAPD分析麻核桃起源与分类地位[J]. 林业科学, 1999, 35(4): 25-30.

WU Y M, PEI D, XI S K. Analysis of the origin and the taxonomic position of Juglans hopeiensis using RAPF markers [J]. Scientia Silvae Sinicae, 1999, 35(4): 25-30.

DANG M, ZHOU H J, WOESTE K E, et al. Comparative phylogeography of Juglans regia and J. mandshurica combining organellar and nuclear DNA markers to assess genetic diversity and introgression in regions of sympatry[J]. Trees, 2021, 35(6): 1993-2007.

OREL G, MARCHANT A D, MCLEOD J A, et al. Characterization of 11 Juglandaceae genotypes based on morphology, cpDNA, and RAPD[J]. HortScience, 2003, 38(6): 1178-1183.

ROGERS R. Temperate Ecosystems|Juglandaceae[M]//Encyclopedia of Forest Sciences. Amsterdam: Elsevier, 2004: 1427-1430.

J∅RGENSEN T, HAILE J, MÖLLER P, et al. A comparative study of ancient sedimentary DNA, pollen and macrofossils from permafrost sediments of northern Siberia reveals long-term vegetational stability[J]. Molecular Ecology, 2012, 21(8): 1989-2003.

JANSSENS S B, KNOX E B, HUYSMANS S, et al. Rapid radiation of Impatiens (Balsaminaceae) during Pliocene and Pleistocene: Result of a global climate change[J]. Molecular Phylogenetics and Evolution, 2009, 52(3): 806-824.

GAO J, WANG B S, MAO J F, et al. Demography and speciation history of the homoploid hybrid pine Pinus densataon the Tibetan Plateau[J]. Molecular Ecology, 2012, 21(19): 4811-4827.

QI X S, CHEN C, COMES H P, et al. Molecular data and ecological niche modelling reveal a highly dynamic evolutionary history of the East Asian Tertiary relict Cercidiphyllum (Cercidiphyllaceae)[J]. New Phytologist, 2012, 196(2): 617-630.

SUN Y W, HOU N, WOESTE K, et al. Population genetic structure and adaptive differentiation of iron walnut Juglans regia subsp. sigillata in southwestern China[J]. Ecology and Evolution, 2019, 9(24): 14154-14166.

ZHANG W P, CAO L, LIN X R, et al. Dead-end hybridization in walnut trees revealed by large-scale genomic sequence data[J]. Molecular Biology and Evolution, 2022, 39(1): msab308.

HU Y H, WOESTE K E, DANG M, et al. The complete chloroplast genome of common walnut (Juglans regia) [J]. Mitochondrial DNA Part B, Resources, 2016, 1(1): 189-190.

ALEXANDER D H, NOVEMBRE J, LANGE K. Fast model-based estimation of ancestry in unrelated individuals[J]. Genome Research, 2009, 19(9): 1655-1664.

LIU J Q, SUN Y S, GE X J, et al. Phylogeographic studies of plants in China: Advances in the past and directions in the future[J]. Journal of Systematics and Evolution, 2012, 50(4): 267-275.

QIU Y X, FU C X, COMES H P, et al. Plant molecular phylogeography in China and adjacent regions: Tracing the genetic imprints of Quaternary climate and environmental change in the world’s most diverse temperate flora[J]. Molecular Phylogenetics and Evolution, 2011, 59(1): 225-244.

ZHANG J B, LI R Q, XIANG X G, et al. Integrated fossil and molecular data reveal the biogeographic diversification of the eastern Asian-eastern North American disjunct hickory genus (Carya Nutt.) [J]. PLoS One, 2013, 8(7): e70449.

PEDERSEN M W, GINOLHAC A, ORLANDO L, et al. A comparative study of ancient environmental DNA to pollen and macrofossils from lake sediments reveals taxonomic overlap and additional plant taxa [J]. Quaternary Science Reviews, 2013, 75: 161-168.

LI Y, SMITH T, SVETLANA P, et al. Paleobiogeography of the lotus plant (Nelumbonaceae: Nelumbo): and its bearing on the paleoclimatic changes[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 2014, 399: 284-293.

GENG F, LEI M, ZHANG N, et al. Demographical complexity within walnut species provides insights into the heterogeneity of geological and climatic fluctuations in East Asia[J/OL]. Journal of Systematics and Evolution. (2024-02-28)[2024-05-10]. https://doi.org/10.1111/jse.13061https://doi.org/10.1111/jse.13061.

WEN J. Evolution of eastern Asian-eastern North American biogeographic disjunctions: A few additional issues[J]. International Journal of Plant Sciences, 2001, 162(S6): S117-S122.

MA Z Y, NIE Z L, LIU X Q, et al. Phylogenetic relationships, hybridization events, and drivers of diversification of East Asian wild grapes as revealed by phylogenomic analyses[J]. Journal of Systematics and Evolution, 2023, 61(2): 273-283.

QIAN H, RICKLEFS R E. Geographical distribution and ecological conservatism of disjunct genera of vascular plants in eastern Asia and eastern North America [J]. Journal of Ecology, 2004, 92(2): 253-265.

XIANG Q Y J, ZHANG W H, RICKLEFS R E, et al. Regional differences in rates of plant speciation and molecularevolution: A comparison between eastern Asia and eastern North America [J]. Evolution, 2004, 58(10): 2175-2184.

DONOGHUE M J, SMITH S A. Patterns in the assembly of temperate forests around the Northern Hemisphere[J]. Philosophical Transactions of the Royal Society of London, Series B, Biological Sciences, 2004, 359(1450): 1633-1644.

HAMILTON J A, ECKERT C G. Population genetic consequences of geographic disjunction: A prairie plant isolated on Great Lakes alvars[J]. Molecular Ecology, 2007, 16(8): 1649-1660.

ZHOU W B, SHI W, SOLTIS P S, et al. Foliar endophyte diversity in Eastern Asian-Eastern North American disjunct tree species-influences of host identity, environment, phylogeny, and geographic isolation[J]. Frontiers in Plant Science, 2023, 14: 1274746.

CHEN J H, HAO Z D, GUANG X M, et al. Liriodendron genome sheds light on angiosperm phylogeny and species-pair differentiation[J]. Nature Plants, 2019, 5(1): 18-25.

COWMAN P F, BELLWOOD D R. Vicariance across major marine biogeographic barriers: Temporal concordance and the relative intensity of hard versus soft barriers[J]. Proceedings Biological Sciences, 2013, 280(1768): 20131541.

KORALL P, PRYER K M. Global biogeography of scaly tree ferns (Cyatheaceae): Evidence for Gondwanan vicariance and limited transoceanic dispersal[J]. Journal of Biogeography, 2014, 41(2): 402-413.

SAURA S, BODIN Ö, FORTIN M J. EDITOR’S CHOICE: Stepping stones are crucial for species’ long-distance dispersal and range expansion through habitat networks[J]. Journal of Applied Ecology, 2014, 51(1): 171-182.

SANZ M, SCHÈNSWETTER P, VALLÈS J, et al. Southern isolation and northern long-distance dispersal shaped the phylogeography of the widespread, but highly disjunct, European high mountain plant Artemisia eriantha (Asteraceae)[J]. Botanical Journal of the Linnean Society, 2014, 174(2): 214-226.

AVISE J C. Phylogeography: The history and formation of species[M]. Cambridge: Harvard University Press, 2000: 230-292.

OTÁLORA M A G, MARTÍNEZ I, ARAGÓN G, et al. Phylogeography and divergence date estimates of a lichen species complex with a disjunct distribution pattern[J]. American Journal of Botany, 2010, 97(2): 216-223.

BIRD C E, FERNANDEZ-SILVA I, SKILLINGS D J, et al. Sympatric speciation in the post “modern synthesis” era of evolutionary biology[J]. Evolutionary Biology, 2012, 39(2): 158180.

RENAUT S, GRASSA C J, YEAMAN S, et al. Genomic islands of divergence are not affected by geography of speciation in sunflowers[J]. Nature Communications, 2013, 4: 1827.

RENAUT S, OWENS G L, RIESEBERG L H. Shared selective pressure and local genomic landscape lead to repeatable patterns of genomic divergence in sunflowers [J]. Molecular Ecology, 2014, 23(2): 311-324.

FITZPATRICK B M, TURELLI M. The geography of mammalian speciation: Mixed signals from phylogenies and range maps[J]. Evolution, 2006, 60(3): 601-615.

KISEL Y, BARRACLOUGH T G. Speciation has a spatial scale that depends on levels of gene flow[J]. The American Naturalist, 2010, 175(3): 316-334.

ZHENG X M, GE S. Ecological divergence in the presence of gene flow in two closely related Oryza species (Oryza rufipogon and O. nivara)[J]. Molecular Ecology, 2010, 19(12): 2439-2454.

BÜSSE S, VON GRUMBKOW P, HUMMEL S, et al. Phylogeographic analysis elucidates the influence of the ice ages on the disjunct distribution of relict dragonflies in Asia[J]. PLoS One, 2012, 7(5): e38132.

GROSSENBACHER D L, VELOZ S D, SEXTON J P. Niche and range size patterns suggest that speciation begins in small, ecologically diverged populations in North American monkeyflowers (Mimulus spp.)[J]. Evolution, 2014, 68(5): 1270-1280.

LI Q Q, KHASBAGAN , ZHANG Z P, et al. Plastid phylogenomics of the tribe potentilleae (Rosaceae)[J]. Molecular Phylogenetics and Evolution, 2024, 190: 107961.

MA Z Y, NIE Z L, REN C, et al. Phylogenomic relationships and character evolution of the grape family (Vitaceae)[J]. Molecular Phylogenetics and Evolution, 2021, 154: 106948.

MA Z Y, WEN J, ICKERT-BOND S M, et al. Phylogenomics, biogeography, and adaptive radiation of grapes[J]. Molecular Phylogenetics and Evolution, 2018, 129: 258-267.

NIE Z L, SUN H, CHEN Z D, et al. Molecular phylogeny and biogeographic diversification of Parthenocissus (Vitaceae) disjunct between Asia and North America[J]. American Journal of Botany, 2010, 97(8): 1342-1353.

WARREN D L, GLOR R E, TURELLI M. Environmental niche equivalency versus conservatism: Quantitative approaches to niche evolution[J]. Evolution, 2008, 62(11): 2868-2883.

EVANS M E K, SMITH S A, FLYNN R S, et al. Climate, niche evolution, and diversification of the ”bird-cage” evening primroses (Oenothera, sections Anogra and Kleinia)[J]. American Naturalist, 2009, 173: 225-240.

DORMANN C F, GRUBER B, WINTER M, et al. Evolution of climate niches in European mammals？[J]. Biology Letters, 2010, 6(2): 229-232.

NAKAZATO T, WARREN D L, MOYLE L C. Ecological and geographic modes of species divergence in wild tomatoes[J]. American Journal of Botany, 2010, 97(4): 680-693.

BROWNSTEIN C D, NEAR T J. Phylogenetics and the Cenozoic radiation of lampreys[J]. Current Biology, 2023, 33(2): 397-404.

BOJKO J, REINKE A W, STENTIFORD G D, et al. Microsporidia: A new taxonomic, evolutionary, and ecological synthesis[J]. Trends in Parasitology, 2022, 38(8): 642-659.

BEATTY G E, PROVAN J. Refugial persistence and postglacial recolonization of North America by the cold-tolerant herbaceous plant Orthilia secunda[J]. Molecular Ecology, 2010, 19(22): 5009-5021.

BEATTY G E, PROVAN J. Phylogeographic analysis of North American populations of the parasitic herbaceous plant Monotropa hypopitys L. reveals a complex history of range expansion from multiple late glacial refugia[J]. Journal of Biogeography, 2011, 38(8): 1585-1599.

BEATTY G E, PROVAN J. Post-glacial dispersal, rather than in situ glacial survival, best explains the disjunct distribution of the Lusitanian plant species Daboecia cantabrica (Ericaceae)[J]. Journal of Biogeography, 2013, 40(2): 335-344.

MORRONE J J. When phylogenetics met biogeography: Willi Hennig, Lars Brundin and the roots of phylogenetic and cladistic biogeography[J]. Cladistics, 2023, 39(1): 58-69.

SKEMA C, JOURDAIN-FIEVET L, DUBUISSON J Y, et al. Out of Madagascar, repeatedly: The phylogenetics and biogeography of Dombeyoideae (Malvaceae s.l.)[J]. Molecular Phylogenetics and Evolution, 2023, 182: 107687.

COWLING R M, LOMBARD A T. Heterogeneity, speciation/extinction history and climate: Explaining regional plant diversity patterns in the Cape Floristic Region[J]. Diversity and Distributions, 2002, 8(3): 163-179.

CARSTENS B C, KNOWLES L L. Shifting distributions and speciation: Species divergence during rapid climate change[J]. Molecular Ecology, 2007, 16(3): 619-627.

HUA X, WIENS J J. How does climate influence speciation？[J]. The American Naturalist, 2013, 182(1): 1-12.

HADID Y, PAVLÍĈEK T, BEILES A, et al. Sympatric incipient speciation of spiny mice Acomys at “Evolution Canyon, ” Israel[J]. Proceedings of the National Academy of Sciences of the United States of America, 2014, 111(3): 1043-1048.

ALSOS I G, EHRICH D, THUILLER W, et al. Genetic consequences of climate change for northern plants[J]. Proceedings Biological Sciences, 2012, 279(1735): 2042-2051.

WANG W, ORTIZ R C, JACQUES F M B, et al. Menispermaceae and the diversification of tropical rainforests near the Cretaceous—Paleogene boundary [J]. New Phytologist, 2012, 195(2): 470-478.

HAMANN E, BLEVINS C, FRANKS S J, et al. Climate change alters plant—herbivore interactions[J]. New Phytologist, 2021, 229(4): 1894-1910.

STEWART J R, STRINGER C B. Human evolution out of Africa: The role of refugia and climate change[J]. Science, 2012, 335(6074): 1317-1321.

STEWART J R, LISTER A M, BARNES I, et al. Refugia revisited: Individualistic responses of species in space and time[J]. Proceedings Biological Sciences, 2010, 277(1682): 661-671.

HEWITT G M. Quaternary phylogeography: The roots of hybrid zones[J]. Genetica, 2011, 139(5): 617-638.

WANG I J, GLOR R E, LOSOS J B. Quantifying the roles of ecology and geography in spatial genetic divergence[J]. Ecology Letters, 2013, 16(2): 175-182.

DAHAL N, KUMAR S, NOON B R, et al. The role of geography, environment, and genetic divergence on the distribution of pikas in the Himalaya[J]. Ecology and Evolution, 2020, 10(3): 1539-1551.

STEWART J R. LISTER A M. Cryptic northern refugia and the origins of the modern biota[J]. Trends in Ecology & Evolution, 2001, 16(11): 608-613.

TZEDAKIS P C, EMERSON B C, HEWITT G M. Cryptic or mystic? Glacial tree refugia in northern Europe[J]. Trends in Ecology & Evolution, 2013, 28(12): 696-704.

ANGELIS K, ÁLVAREZ-CARRETERO S, DOS REIS M, et al. An evaluation of different partitioning strategies for Bayesian estimation of species divergence times[J]. Systematic Biology, 2018, 67(1): 61-77.

TEJADA J V, ANTOINE P O, MÜNCH P, et al. Bayesian total-evidence dating revisits sloth phylogeny and biogeography: A cautionary tale on morphological clock analyses[J]. Systematic Biology, 2024, 73(1): 125-139.

LINDSEY C R, KNOLL A H, HERRON M D, et al. Fossil-calibrated molecular clock data enable reconstruction of steps leading to differentiated multicellularity and anisogamy in the Volvocine algae[J]. BMC Biology, 2024, 22(1): 79.

JIA D R, ABBOTT R J, LIU T L, et al. Out of the Qinghai-Tibet Plateau: Evidence for the origin and dispersal of Eurasian temperate plants from a phylogeographic study of Hippophaë rhamnoides (Elaeagnaceae)[J]. New Phytologist, 2012, 194(4): 1123-1133.

ZHANG G Y, SONG Y T, CHEN N, et al. Chromosome-level genome assembly of Hippophae tibetana provides insights into high-altitude adaptation and flavonoid biosynthesis[J]. BMC Biology, 2024, 22(1): 82.

HOBAN S M, BORKOWSKI D S, BROSI S L, et al. Range-wide distribution of genetic diversity in the North American tree Juglans cinerea: A product of range shifts, not ecological marginality or recent population decline[J]. Molecular Ecology, 2010, 19(22): 4876-4891.

WANG W T, XU B, ZHANG D Y, et al. Phylogeography of postglacial range expansion in Juglans mandshurica (Juglandaceae) reveals no evidence of bottleneck, loss of genetic diversity, or isolation by distance in the leading-edge populations[J]. Molecular Phylogenetics and Evolution, 2016, 102: 255-264.

HUANG W P, SUN H, DENG T, et al. Molecular phylogenetics and biogeography of the eastern Asian-eastern North American disjunct Mitchella and its close relative Damnacanthus (Rubiaceae, Mitchelleae)[J]. Botanical Journal of the Linnean Society, 2013, 171(2): 395-412.

FICHANT T, LEDENT A, COLLART F, et al. Dispersal capacities of pollen, seeds and spores: Insights from comparative analyses of spatial genetic structures in bryophytes and spermatophytes[J]. Frontiers in Plant Science, 2023, 14: 1289240.

OMONDI S F, GITHAE E W, KHASA D P. Long-distance gene flow in Acacia senegal: Hope for disturbed and fragmented populations[J]. Ecology and Evolution, 2023, 13(7): e10292.

LI E Z, WANG Y S, LIU K J, et al. Historical climate change and vicariance events contributed to the intercontinental disjunct distribution pattern of ash species (Fraxinus, Oleaceae)[J]. Communications Biology, 2024, 7(1): 603.

LUO X, ZHOU H J, CAO D, et al. Domestication and selection footprints in Persian walnuts (Juglans regia)[J]. PLoS Genetics, 2022, 18(12): e1010513.

HAN H, WOESTE K E, HU Y H, et al. Genetic diversity and population structure of common walnut (Juglans regia) in China based on EST-SSRs and the nuclear gene phenylalanine ammonia-lyase (PAL)[J]. Tree Genetics & Genomes, 2016, 12(6): 111.

DANG M, YUE M, ZHANG M, et al. Gene introgression among closely related species in sympatric populations: A case study of three walnut (Juglans) species[J]. Forests, 2019, 10(11): 965.

MANNI F, GUÉRARD E, HEYER E. Geographic patterns of (genetic, morphologic, linguistic) variation: How barriers can be detected by using Monmonier’s algorithm[J]. Human Biology, 2004, 76(2): 173-190.

DARWIN C. On the origin of species by means of natural selection: Or, the preservation of favoured races in the struggle for life[M]. London: John Murray, 1859: 7-19.

HOSKIN C J, HIGGIE M, MCDONALD K R, et al. Reinforcement drives rapid allopatric speciation[J]. Nature, 2005, 437(7063): 1353-1356.

APRIL J, HANNER R H, DION-CÔTÉ A M, et al. Glacial cycles as an allopatric speciation pump in northeastern American freshwater fishes[J]. Molecular ecology, 2013, 22(2): 409-422.

BECKER F S, ALEXANDER G J, TOLLEY K A. Substrate specialisation drives an unexpectedly diverse radiation in barking geckos (Ptenopus: Gekkonidae)[J]. Molecular Phylogenetics and Evolution, 2024, 197: 108104.

GAVRILETS S. Perspective: Models of speciation: What have we learned in 40 years？[J]. Evolution, 2003, 57(10): 2197-2215.

XIAO L Q, MÖLLER M, ZHU H. High nrDNA ITS polymorphism in the ancient extant seed plant Cycas: Incomplete concerted evolution and the origin of pseudogenes[J]. Molecular Phylogenetics and Evolution, 2010, 55(1): 168-177.

DUPIN J, HONG-WA C, GAUDEUL M, et al. Phylogenetics and biogeography of the olive family (Oleaceae)[J]. Annals of Botany, 2024: mcae100.

ABBOTT R, ALBACH D, ANSELL S, et al. Hybridization and speciation[J]. Journal of Evolutionary Biology, 2013, 26(2): 229-246.

PHILLIPSEN I C, KIRK E H, BOGAN M T, et al. Dispersal ability and habitat requirements determine landscape-level genetic patterns in desert aquatic insects[J]. Molecular Ecology, 2015, 24(1): 54-69.

BANK C, BÜRGER R, HERMISSON J. The limits to parapatric speciation: Dobzhansky-Muller incompatibilities in a continent-island model[J]. Genetics, 2012, 191(3): 845-863.

DIECKMANN U, DOEBELI M. On the origin of species by sympatric speciation[J]. Nature, 1999, 400(6742): 354-357.

BARLUENGA M, STÖLTING K N, SALZBURGER W, et al. Sympatric speciation in Nicaraguan crater lake cichlid fish[J]. Nature, 2006, 439(7077): 719-723.

NOSIL P. Speciation with gene flow could be common[J]. Molecular Ecology, 2008, 17(9): 21032106.

SCHMID S, BACHMANN SALVY M, GARCIA JIMENEZ A, et al. Gene flow throughout the evolutionary history of a colour polymorphic and generalist clownfish[J]. Molecular Ecology, 2024, 33(14): e17436.

LIU J, NIE Z L, REN C, et al. Phylogenomics of Aralia sect. Aralia (Araliaceae): Signals of hybridization and insights into its species delimitations and intercontinental biogeography[J]. Molecular Phylogenetics and Evolution, 2023, 181: 107727.

BOLTE C E, PHANNARETH T, ZAVALA-PAEZ M, et al. Genomic insights into hybrid zone formation: The role of climate, landscape, and demography in the emergence of a novel hybrid lineage[J]. Molecular Ecology, 2024, 33(14): e17430.

ROSSER N, SEIXAS F, QUESTE LM, et al. Hybrid speciation driven by multilocus introgression of ecological traits[J]. Nature, 2024, 628(8009): 811-817.

WIESE J. Digest: Pelagic habitats promote speciation but constrain morphological evolution[J]. Evolution, 2024: qpae091.

NOSIL P. Ecological speciation[M]. Oxford: Oxford University Press, 2012: 280.

SCHLUTER D. Evidence for ecological speciation and its alternative[J]. Science, 2009, 323(5915): 737-741.

SCHLUTER D, CONTE G L. Genetics and ecological speciation[J]. Proceedings of the National Academy of Sciences of the United States of America, 2009, 106(S1): 9955-9962.

SOBEL J M, CHEN G F, WATT L R, et al. The biology of speciation[J]. Evolution, 2010, 64(2): 295-315.

MAYA-LASTRA C A, SWEENEY P W, EATON D A R, et al. Caught in the act: Incipient speciation at the southern limit of viburnum in the central Andes[J]. Systematic Biology, 2024: syae023.

LI Y R, FRITSCH P W, ZHAO G G, et al. Population differentiation and dynamics of five pioneer species of Gaultheria from the secondary forests in subtropical China[J]. BMC Plant Biology, 2024, 24(1): 516.

SUN P W, CHANG J T, LUO M X, et al. Genomic insights into local adaptation and vulnerability of Quercus longinux to climate change[J]. BMC Plant Biology, 2024, 24(1): 279.

CIANCHI R, ARDUINO P, MOSCO M C, et al. Evidence of hybrid speciation in the North American primroses Primula suffrutescensP. parryiP. rusbyi and P. angustifolia (Primulaceae)[J]. Plant Biosystems-an International Journal Dealing with All Aspects of Plant Biology, 2015, 149(2): 229-234.

ROSSETTO M, ALLEN C B, THURLBY K A G, et al. Genetic structure and bio-climatic modeling support allopatric over parapatric speciation along a latitudinal gradient[J]. BMC Evolutionary Biology, 2012, 12: 149.

SEXTON J P, HANGARTNER S B, HOFFMANN A A. Genetic isolation by environment or distance: Which pattern of gene flow is most common? [J]. Evolution, 2014, 68(1): 1-15.

FRISTOE T S, BLEILEVENS J, KINLOCK N L, et al. Evolutionary imbalance, climate and human history jointly shape the global biogeography of alien plants[J]. Nature Ecology & Evolution, 2023, 7(10): 1633-1644.

BROWN L E, KHAMIS K, WILKES M, et al. Functional diversity and community assembly of river invertebrates show globally consistent responses to decreasing glacier cover[J]. Nature Ecology & Evolution, 2018, 2(2): 325-333.

LI J, ZHENG Z, HUANG K Y, et al. Vegetation changes during the past 40, 000 years in Central China from a long fossil record[J]. Quaternary International, 2013, 310: 221-226.

SMITH T, ROSE K D, GINGERICH P D. Rapid Asia-Europe-North America geographic dispersal of earliest Eocene primate Teilhardina during the Paleocene-Eocene Thermal Maximum[J]. Proceedings of the National Academy of Sciences of the United States of America, 2006, 103(30): 11223-11227.

STANKOWSKI S. Ecological speciation in an island snail: Evidence for the parallel evolution of a novel ecotype and maintenance by ecologically dependent postzygotic isolation[J]. Molecular Ecology, 2013, 22(10): 2726-2741.

SHAFER A B A, WOLF J B W. Widespread evidence for incipient ecological speciation: A meta-analysis of isolation-by-ecology[J]. Ecology Letters, 2013, 16(7): 940-950.

RÄSÄNEN K, HENDRY A P. Disentangling interactions between adaptive divergence and gene flow when ecology drives diversification[J]. Ecology Letters, 2008, 11(6): 624-636.

SERVEDIO M R, HERMISSON J, VAN DOORN G S. Hybridization may rarely promote speciation[J]. Journal of Evolutionary Biology, 2013, 26(2): 282-285.

NOSIL P, VINES T H, FUNK D J. Perspective: Reproductive isolation caused by natural selection against immigrants from divergent habitats[J]. Evolution, 2005, 59(4): 705-719.

BOLNICK D I, NOSIL P. Natural selection in populations subject to a migration load[J]. Evolution, 2007, 61(9): 2229-2243.

BRIDLE J R, POLECHOVÁ J, KAWATA M, et al. Why is adaptation prevented at ecological margins? New insights from individual-based simulations[J]. Ecology Letters, 2010, 13(4): 485-494.

KARIMI N, KRIEG C P, SPALINK D, et al. Chromosomal evolution, environmental heterogeneity, and migration drive spatial patterns of species richness in Calochortus(Liliaceae)[J]. Proceedings of the National Academy of Sciences of the United States of America, 2024, 121(10): e2305228121.

PLATH M, PFENNINGER M, LERP H, et al. Genetic differentiation and selection against migrants in evolutionarily replicated extreme environments[J]. Evolution, 2013, 67(9): 2647-2661.

GARROWAY C J, RADERSMA R, SEPIL I, et al. Fine-scale genetic structure in a wild bird population: The role of limited dispersal and environmentally based selection as causal factors[J]. Evolution, 2013, 67(12): 3488-3500.

FRANKHAM R, BALLOU J D, ELDRIDGE M D B, et al. Predicting the probability of outbreeding depression[J]. Conservation Biology, 2011, 25(3): 465-475.

PEKKALA N, KNOTT K E, KOTIAHO J S, et al. The benefits of interpopulation hybridization diminish with increasing divergence of small populations[J]. Journal of Evolutionary Biology, 2012, 25(11): 2181-2183.

SOULARUE J P, KREMER A. Assortative mating and gene flow generate clinal phenological variation in trees[J]. BMC Evolutionary Biology, 2012, 12: 79.

BAI W N, YAN P C, ZHANG B W, et al. Demographically idiosyncratic responses to climate change and rapid Pleistocene diversification of the walnut genus Juglans (Juglandaceae) revealed by whole-genome sequences[J]. The New Phytologist, 2018, 217(4): 1726-1736.

BAI W N, ZENG Y F, LIAO W J, et al. Flowering phenology and wind-pollination efficacy of heterodichogamous Juglans mandshurica (Juglandaceae)[J]. Annals of Botany, 2006, 98(2): 397-402.

AVNI R, NAVE M, BARAD O, et al. Wild emmer genome architecture and diversity elucidate wheat evolution and domestication[J]. Science, 2017, 357(6346): 93-97.

DIAMOND J. Evolution, consequences and future of plant and animal domestication[J]. Nature, 2002, 418(6898): 700-707.

GAUT B S, SEYMOUR D K, LIU Q P, et al. Demography and its effects on genomic variation in crop domestication[J]. Nature Plants, 2018, 4(8): 512-520.

MEYER R S, DUVAL A E, JENSEN H R. Patterns and processes in crop domestication: An historical review and quantitative analysis of 203 global food crops[J]. The New Phytologist, 2012; 196(1): 29-48.

HUANG X H, HUANG S W, HAN B, et al. The integrated genomics of crop domestication and breeding[J]. Cell, 2022, 185(15): 2828-2839.

CHENG S F, FENG C, WINGEN L U, et al. Harnessing landrace diversity empowers wheat breeding[J]. Nature, 2024, 632: 823-831.

MEYER R S, PURUGGANAN M D. Evolution of crop species: Genetics of domestication and diversification[J]. Nature Reviews Genetics, 2013, 14(12): 840-852.

XIAO J, LIU B, YAO Y Y, et al. Wheat genomic study for genetic improvement of traits in China[J]. Science China Life Sciences, 2022, 65(9): 1718-1775.

HAAS M, SCHREIBER M, MASCHER M. Domestication and crop evolution of wheat and barley: Genes, genomics, and future directions[J]. Journal of Integrative Plant Biology, 2019, 61(3): 204-225.

FORNASIERO A, WING R A, RONALD P. Rice domestication[J]. Current Biology, 2022, 32(1): R20-R24.

IZAWA T. Reloading DNA history in rice domestication[J]. Plant & Cell Physiology, 2022, 63(11): 1529-1539.

ZHANG J P, JIANG L P, YU L P, et al. Rice’s trajectory from wild to domesticated in East Asia[J]. Science, 2024, 384(6698): 901-906.

YU H, LIN T, MENG X B, et al. A route to de novo domestication of wild allotetraploid rice[J]. Cell, 2021, 184(5): 1156-1170.

STITZER M C, ROSS-IBARRA J. Maize domestication and gene interaction[J]. The New Phytologist, 2018, 220(2): 395-408.

DONG Z B, ALEXANDER M, CHUCK G. Understanding grass domestication through maize mutants[J]. The Trends in Genetics, 2019, 35(2): 118-128.

ABRAHAM-JUÁREZ M J, BARNES A C, ARAGÓN-RAYGOZA A, et al. The arches and spandrels of maize domestication, adaptation, and improvement[J]. Current Opinion in Plant Biology, 2021, 64: 102124.

CHEN Q Y, LI W Y, TAN L B, et al. Harnessing knowledge from maize and rice domestication for new crop breeding[J]. Molecular Plant, 2021, 14(1): 9-26.

ZHU G T, WANG S C, HUANG Z J, et al. Rewiring of the fruit metabolome in tomato breeding[J]. Cell, 2018, 172(1/2): 249-261.

CONSORTIUM T G. The tomato genome sequence provides insights into fleshy fruit evolution[J]. Nature, 2012, 485(7400): 635-641.

CHE G, ZHANG X L. Molecular basis of cucumber fruit domestication[J]. Current Opinion in Plant Biology, 2019, 47: 38-46.

SEDIVY E J, WU F Q, HANZAWA Y. Soybean domestication: The origin, genetic architecture and molecular bases[J]. The New Phytologist, 2017, 214(2): 539-553.

GAO L, GONDA I, SUN H H, et al. The tomato pan-genome uncovers new genes and a rare allele regulating fruit flavor[J]. Nature Genetics, 2019, 51(6): 1044-1051.

UNVER T, WU Z Y, STERCK L, et al. Genome of wild olive and the evolution of oil biosynthesis[J]. Proceedings of the National Academy of Sciences of the United States of America, 2017, 144(44): E9413-E9422.

WU J, WANG Y T, XU J B, et al. Diversification and independent domestication of Asian and European pears[J]. Genome Biology, 2018, 19(1): 77.

LI Y, CAO K, ZHU G R, et al. Genomic analyses of an extensive collection of wild and cultivated accessions provide new insights into peach breeding history[J]. Genome Biology, 2019, 20(1): 36.

BERNARD A, LHEUREUX F, DIRLEWANGER E. Walnut: Past and future of genetic improvement[J]. Tree Genetics & Genomes, 2017, 14(1): 1.

DING Y M, CAO Y, ZHANG W P, et al. Population-genomic analyses reveal bottlenecks and asymmetric introgression from Persian into iron walnut during domestication[J]. Genome Biology, 2022, 23(1): 145.

MARTÍNEZ-GARCÍA P J, CREPEAU M W, PUIU D, et al. The walnut (Juglans regia) genome sequence reveals diversity in genes coding for the biosynthesis of non-structural polyphenols[J]. The Plant Journal, 2016, 87(5): 507-532.

CHEN L N, MA Q G, CHEN Y K, et al. Identification of major walnut cultivars grown in China based on nut phenotypes and SSR markers[J]. Scientia Horticulturae, 2014, 168: 240248.

JI F Y, MA Q G, ZHANG W T, et al. A genome variation map provides insights into the genetics of walnut adaptation and agronomic traits[J]. Genome Biology, 2021, 22(1): 300.

YANG C J, SAMAYOA L F, BRADBURY P J, et al. The genetic architecture of teosinte catalyzed and constrained maize domestication[J]. Proceedings of the National Academy of Sciences of the United States of America, 2019, 116(12): 5643-5652.

HUANG X H, KURATA N, WEI X H, et al. A map of rice genome variation reveals the origin of cultivated rice[J]. Nature, 2012, 490(7421): 497-501.

WU J, WANG L F, FU J J, et al. Resequencing of 683 common bean genotypes identifies yield component trait associations across a north-south cline[J]. Nature Genetics, 2020, 52(1): 118-125.

CLARK R M, WAGLER T N, QUIJADA P, et al. A distant upstream enhancer at the maize domestication gene Tb1 has pleiotropic effects on plant and inflorescent architecture[J]. Nature Genetics, 2006, 38(5): 594-597.

WU G A, TEROL J, IBANEZ V, et al. Genomics of the origin and evolution of Citrus[J]. Nature, 2018, 554(7692): 311-316.

WANG L, HE F, HUANG Y, et al. Genome of wild mandarin and domestication history of mandarin[J]. Molecular Plant, 2018, 11(8): 1024-1037.

ZHOU Y F, MASSONNET M, SANJAK J S, et al. Evolutionary genomics of grape (Vitis vinifera ssp. vinifera) domestication[J]. Proceedings of the National Academy of Sciences of the United States of America, 2017, 114(44): 11715-11720.

ALLABY R G. Two domestications for grapes[J]. Science, 2023, 379(6635): 880-881.

DIEZ C M, TRUJILLO I, MARTINEZ-URDIROZ N, et al. Olive domestication and diversification in the Mediterranean Basin[J]. The New Phytologist, 2015, 206(1): 436-447.

GAUT B S, DÍEZ C M, MORRELL P L. Genomics and the contrasting dynamics of annual and perennial domestication[J]. Trends in Genetisc, 2015, 31(12): 709-719.

MCCLURE K A, SAWLER J, GARDNER K M, et al. Genomics: A potential panacea for the perennial problem[J]. American Journal of Botany, 2014, 101(10): 1780-1790.

MILLER A J, GROSS B L. From forest to field: Perennial fruit crop domestication[J]. American Journal of Botany, 2011, 98(9): 1389-1414.

HARFOUCHE A, MEILAN R, KIRST M, et al. Accelerating the domestication of forest trees in a changing world[J]. Trends in Plant Science, 2012, 17(2): 64-72.

BAYAZIT S, KAZAN K, GÜLBITTI S, et al. AFLP analysis of genetic diversity in low chill requiring walnut (Juglans regia L.) genotypes from Hatay, Turkey[J]. Scientia Horticulturae, 2007, 111(4): 394-398.

ZOHARY D, HOPF M, WEISS E. Domestication of plants in the Old World: The origin and spread of domesticated plants in south-west Asia, Europe, and the Mediterranean Basin[M]. 4th ed. Oxford: Oxford University Press, 2012: 1-68.

ROOR W, KONRAD H, MAMADJANOV D, et al. Population differentiation in common walnut (Juglans regia L.) across major parts of its native range—Insights from molecular and morphometric data[J]. Journal of Heredity, 2017, 108(4): 391-404.

ZHANG B W, XU L L, LI N, et al. Phylogenomics reveals an ancient hybrid origin of the Persian walnut [J]. Molecular Biology and Evolution, 2019, 36(11): 2451-2461.

WANG J T, YE H, ZHOU H J, et al. Genome-wide association analysis of 101 accessions dissects the genetic basis of shell thickness for genetic improvement in Persian walnut (Juglans regia L.)[J]. BMC Plant Biology, 2022, 22(1): 436.

SONG B, NING W D, WEI D, et al. Plant genome resequencing and population genomics: Current status and future prospects[J]. Molecular Plant, 2023, 16(8): 1252-1268.

LONG Q M, CAO S, HUANG G Z, et al. Population comparative genomics discovers gene gain and loss during grapevine domestication[J]. Plant Physiology, 2024, 195(2): 1401-1413.

MARRANO A, BRITTON M, ZAINI P A, et al. High-quality chromosome-scale assembly of the walnut (Juglans regia L.) reference genome[J]. GigaScience, 2020, 9(5): giaa050.

STEVENS K A, WOESTE K, CHAKRABORTY S, et al. Genomic variation among and within six Juglans species[J]. G3: Genes, Genomes, Genetics, 2018, 8(7): 2153-2165.

ZHU T T, WANG L, YOU F M, et al. Sequencing a Juglans regia×J. microcarpa hybrid yields high-quality genome assemblies of parental species[J]. Horticulture Research, 2019, 6: 55.

ZHANG J P, ZHANG W T, JI F Y, et al. A high-quality walnut genome assembly reveals extensive gene expression divergences after whole-genome duplication[J]. Plant Biotechnology Journal, 2020, 18(9): 1848-1850.

NING D L, WU T, XIAO L J, et al. Chromosomal-level assembly of Juglans sigillata genome using Nanopore, BioNano, and Hi-C analysis[J]. GigaScience, 2020, 9(2): giaa006.

WANG Y, YANG Y X, YUAN X L, et al. Draft genome sequence of endophytic fungus Talaromyces purpureogenuss train YAFEF302, isolated from Juglans sigillata[J]. Microbiology Resource Announcements, 2024, 13(1): e0082923.

YAN F, XI R M, SHE R X, et al. Improved de novo chromosome-level genome assembly of the vulnerable walnut tree Juglans mandshurica reveals gene family evolution and possible genome basis of resistance to lesion nematode[J]. Molecular Ecology Resources, 2021, 21(6): 2063-2076.

LI X, CAI K W, ZHANG Q H, et al. The manchurian walnut genome: Insights into juglone and lipid biosynthesis[J]. GigaScience, 2022, 11: giac057.

ZHOU H J, YAN F, HAO F, et al. Pan-genome and transcriptome analyses provide insights into genomic variation and differential gene expression profiles related to disease resistance and fatty acid biosynthesis in eastern black walnut (Juglans nigra)[J]. Horticulture Research, 2023, 10(3): uhad015.

FITZ-GIBBON S, MEAD A, O’DONNELL S, et al. Reference genome of California walnut, Juglans californica, and resemblance with other genomes in the order Fagales[J]. The Journal of Heredity, 2023, 114(5): 570-579.

HAN L Q, LUO X, ZHAO Y, et al. A haplotype-resolved genome provides insight into allele-specific expression in wild walnut (Juglans regia L.)[J]. Scientific Data, 2024, 11(1): 278.

GUZMAN-TORRES C R, TRYBULEC E, LEVASSEUR H, et al. Conserving a threatened North American walnut: A chromosome-scale reference genome for butternut (Juglans cinerea)[J]. G3: Genes, Genomes, Genetics, 2024, 14(2): jkad189.

QU Y Q, SHANG X L, ZENG Z Y, et al. Whole-genome duplication reshaped adaptive evolution in a relict plant species, Cyclocarya paliurus[J]. Genomics, Proteomics & Bioinformatics, 2023, 21(3): 455-469.

QU Y Q, SHANG X L, FANG S Z, et al. Genome assembly of two diploid and one auto-tetraploid Cyclocarya paliurus genomes[J]. Scientific Data, 2023, 10(1): 507.

YU R M, ZHANG N, ZHANG B W, et al. Genomic insights into biased allele loss and increased gene numbers after genome duplication in autotetraploid Cyclocarya paliurus[J]. BMC Biology, 2023, 21(1): 168.

DING Y M, PANG X X, CAO Y, et al. Genome structure-based Juglandaceae phylogenies contradict alignment-based phylogenies and substitution rates vary with DNA repair genes[J]. Nature Communications, 2023, 14(1): 617.

CAO Y, ALMEIDA-SILVA F, ZHANG W P, et al. Genomic insights into adaptation to karst limestone and incipient speciation in East Asian Platycarya spp. (Juglandaceae)[J]. Molecular Biology and Evolution, 2023, 40(6): msad121.

ZHOU H J, ZHANG X D, LIU H Z, et al. Chromosome-level genome assembly of Platycarya strobilacea[J]. Scientific Data, 2024, 11(1): 269.

LIU H Z, ZHOU H T, YE H, et al. Integrated metabolomic and transcriptomic dynamic profiles of endopleura coloration during fruit maturation in three walnut cultivars[J]. BMC Plant Biology, 2024, 24(1): 109.

MA J Y, ZUO D J, YE H, et al. Genome-wide identification, characterization, and expression pattern of the late embryogenesis abundant (LEA) gene family in Juglans regia and its wild relatives J. mandshurica[J]. BMC Plant Biology, 2023, 23(1): 80.

MA J Y, ZUO D J, ZHANG X D, et al. Genome-wide identification analysis of the 4-Coumarate: CoA ligase (4CL) gene family expression profiles in Juglans regia and its wild relatives J. Mandshurica resistance and salt stress[J]. BMC Plant Biology, 2024, 24(1): 211.

LI M D, OU M W, HE X Z, et al. DNA methylation role in subgenome expression dominance of Juglans regia and its wild relative J. mandshurica[J]. Plant Physiology, 2023, 193(2): 1313-1329.

ZHOU H J, MA J Y, LIU H Z, et al. Genome-wide identification of the CBF gene family and ICE transcription factors in walnuts and expression profiles under cold conditions[J]. International Journal of Molecular Science, 2023, 25(1): 25.

CHEN F, CHEN J, WANG Z, et al. Genomics: Cracking the mysteries of walnuts[J]. Jounal of Genetics, 2019, 98(2): 33.

CHEN M J, FAN W J, JI F Y, et al. Genome-wide identification of agronomically important genes in outcrossing crops using OutcrossSeq[J]. Molecular Plant, 2021, 14(4): 556-570.

GUILLAUME C, ISABELLE C, MARC B, et al. Assessing frost damages using dynamic models in walnut trees: Exposure rather than vulnerability controls frost risks[J]. Plant, Cell & Environment, 2018, 41(5): 1008-1021.

SONG J M, GUAN Z L, HU J L, et al. Eight high-quality genomes reveal pan-genome architecture and ecotype differentiation of Brassica napus[J]. Nature Plants, 2020, 6(1): 34-45.

ZHOU R, DONG Y H, LIU X, et al. JrWRKY21 interacts with JrPTI5L to activate the expression of JrPR5L for resistance to Colletotrichum gloeosporioides in walnut[J]. The Plant Journal, 2022, 111(4): 1152-1166.

ARAB M M, BROWN P J, ABDOLLAHI-ARPANAHI R, et al. Genome-wide association analysis and pathway enrichment provide insights into the genetic basis of photosynthetic responses to drought stress in Persian walnut[J]. Horticulture Research, 2022, 9: uhac124.

ARAB M M, MARRANO A, ABDOLLAHI-ARPANAHI R, et al. Combining phenotype, genotype, and environment to uncover genetic components underlying water use efficiency in Persian walnut[J]. Journal of Experimental Botany, 2020, 71(3): 1107-1127.